I have had a long-standing interest in the control of behavior, initially examining how circadian clocks organize behavior and how peptide and steroid hormones initiate and direct behavioral programs.   I have also been fascinated by metamorphosis, especially how it evolved within the insects and how the nervous system and behavior deals with the transition from larva to adult.  The latter interest has led to research on the remodeling of larval neurons and circuits during metamorphosis and on the production of lineages of adult-specific neurons during larval growth and their eventual incorporation into the mature CNS.   The adult-specific neurons are the focus of current interests in understanding the functional roles of the neurons that arise in each lineage and how these lineages have expanded and diversified to support behavioral innovations during the evolution of the nervous system.


B.S.  (Biology) University  of Notre Dame, 1967.
Ph.D. (Biology) Harvard University, 1970.
Junior Fellow, Harvard Society of Fellows, 1970-73.
Assistant Professor of Zoology, University of Washington, 1973-76.
Associate Professor of Zoology, University of Washington, 1976-78.
Professor of Zoology, University of Washington, 1978-2003
Professor of Biology, University of Washington, 2003 – 2007
Professor Emeritus, University of Washington, 2007 – date
Group Leader, HHMI Janelia Research Campus,  2007 – 2014
Senior Fellow, HHMI Janelia Research Campus, 2014 – date.


NSF Predoctoral Fellowship (1967-70)
Harvard Junior Fellow (1970-73)
A.A.A.S. Newcomb Cleveland Research Prize (1970)
NIH Research Career Development Award (1976-81)
McKnight Foundation Neurosciences Development Award (1981-84)
John Simon Guggenheim Fellowship (1986-87)
NIH Jacob K. Javits Award (1988-1995)
Entomological Soc. America Founder's Award (1990)
Foreign Fellow, Churchill College, Cambridge University (2000)
Wigglesworth Metal in Insect Biology, 2008
Fellow, American Academy of Arts and Sciences, 2009


296 research articles and reviews in the areas of invertebrate behavior, development, neurobiology and endocrinology.